The airline makes a sizeable settlement offer to the surviving Seattle Grace docs for the horrific plane crash that took the lives of two of their own. The decision to take the money has to be unanimous. Cristina tells Owen that she'll do whatever everyone else wants to do. This is discussed during one of the many phone chats they've been having lately. Meredith and Callie, who is serving as proxy for Mark and Arizona, seem to be leaning toward settlement. Derek, however, is struggling with some internal conflict. More on that later.
Everyone is shocked to see that April is back, especially Jackson. The two of them team up on a case involving the skin removal of a woman who lost a significant amount of weight. Things are awkward from the get-go since April announces that she's re-virginizing. In other words, she's acting like what she went through with Jackson (we'll call it her "de-virginizing) never happened. Meredith figures out their secret and realizes she missed a lot during that plane crash like the fact that Alex is diseased man-whore again.
Speaking of Alex, he's warned by Callie to stop sleeping with interns because it ruins them. Guess that's why he's being so mean to his latest pretty underling, Jo, even though he totally digs chicks with boy names. Alex puts the kibosh on Jo's hopes of scrubbing in the surgery of a young girl with a nearly severed foot. As for the patient, she wants to get back to her solo sailing trip. Callie warns her way-to-passive parents to stop her even if means having their daughter hate them. Oh, and Alex eventually does let Jo scrub in for a surgery.